About the Authors

Candace K. Mathiason

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Sheila A. Hays

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Jenny Powers

Affiliation National Park Service, Fort Collins, Colorado, United States of America

Jeanette Hayes-Klug

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Julia Langenberg

Affiliation Wisconsin Department of Natural Resources, Madison, Wisconsin, United States of America

Sallie J. Dahmes

Affiliation WASCO Inc., Monroe, Georgia, United States of America

David A. Osborn

Affiliation Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

Karl V. Miller

Affiliation Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

Robert J. Warren

Affiliation Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, United States of America

Gary L. Mason

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Edward A. Hoover

Edward.Hoover@ColoState.edu

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CKM EAH. Performed the experiments: CKM SAH JGP JHK. Analyzed the data: CKM GLM. Contributed reagents/materials/analysis tools: JL SJD DAO KVM RJW. Wrote the paper: CKM. Paper editing: EAH JGP JL.