About the Authors

Katherine L. Hammond

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Helen E. Loynes

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Catriona Mowbray

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Greg Runke

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Matthias Hammerschmidt

Current address: Institute for Developmental Biology, Cologne University, Koeln, Germany

Affiliation Max-Planck-Institute of Immunobiology, Freiburg, Germany

Mary C. Mullins

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Victoria Hildreth

Affiliation Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, United Kingdom

Bill Chaudhry

Affiliation Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, United Kingdom

Tanya T. Whitfield

t.whitfield@sheffield.ac.uk

Affiliation MRC Centre for Developmental and Biomedical Genetics and Department of Biomedical Science, University of Sheffield, Sheffield, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KLH MH MM BC TW. Performed the experiments: KLH HEL CM GR MH MM. Analyzed the data: KLH HEL CM VH BC TW. Contributed reagents/materials/analysis tools: MH MM BC. Wrote the paper: KLH TW.