About the Authors

Jonathan H. Lin

JLin@ucsd.edu

Affiliations Howard Hughes Medical Institute, University of California San Francisco, San Francisco, California, United States of America, Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Han Li

Affiliations Howard Hughes Medical Institute, University of California San Francisco, San Francisco, California, United States of America, Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Yuhong Zhang

Affiliation Departments of Cell Biology and Medicine, Kimmel Center for Biology and Medicine, Skirball Institute, New York University School of Medicine, New York, New York, United States of America

David Ron

Affiliation Departments of Cell Biology and Medicine, Kimmel Center for Biology and Medicine, Skirball Institute, New York University School of Medicine, New York, New York, United States of America

Peter Walter

Affiliations Howard Hughes Medical Institute, University of California San Francisco, San Francisco, California, United States of America, Department of Biochemistry and Biophysics, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JHL HL YZ DR PW. Performed the experiments: JHL HL YZ. Analyzed the data: JHL HL YZ DR PW. Contributed reagents/materials/analysis tools: JHL YZ DR PW. Wrote the paper: JHL PW.