About the Authors

John P. Swaddle

jpswad@wm.edu

Affiliations National Center for Ecological Analysis and Synthesis, University of California Santa Barbara, Santa Barbara, California, United States of America, Institute of Integrative Bird Behavior Studies, Biology Department, College of William and Mary, Williamsburg, Virginia, United States of America

Stavros E. Calos

Affiliation Institute of Integrative Bird Behavior Studies, Biology Department, College of William and Mary, Williamsburg, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JS. Performed the experiments: JS SC. Analyzed the data: JS SC. Contributed reagents/materials/analysis tools: JS. Wrote the paper: JS SC.