About the Authors

Necat Havlioglu

Contributed equally to this work with: Necat Havlioglu, Jun Wang

Affiliation Department of Pathology, Saint Louis University, St. Louis, Missouri, United States of America

Jun Wang

Contributed equally to this work with: Necat Havlioglu, Jun Wang

Affiliation Department of Ecology and Evolution, The University of Chicago, Chicago, Illinois, United States of America

Kazuo Fushimi

Affiliation Department of Neurology, Lurie Comprehensive Cancer Center, Center for Genetic Medicine, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Maria D. Vibranovski

Affiliation Department of Ecology and Evolution, The University of Chicago, Chicago, Illinois, United States of America

Zhengyan Kan

Affiliation Department of Genetics, Washington University in St. Louis, St. Louis, Missouri, United States of America

Warren Gish

Affiliation Department of Genetics, Washington University in St. Louis, St. Louis, Missouri, United States of America

Alexei Fedorov

Affiliation Department of Medicine and Program in Bioinformatics and Proteomics/Genomics, Medical University of Ohio, Toledo, Ohio, United States of America

Manyuan Long

To whom correspondence should be addressed. E-mail: mlong@midway.uchicago.edu (ML); jane-wu@northwestern.edu (JW)

Affiliation Department of Ecology and Evolution, The University of Chicago, Chicago, Illinois, United States of America

Jane Y. Wu

To whom correspondence should be addressed. E-mail: mlong@midway.uchicago.edu (ML); jane-wu@northwestern.edu (JW)

Affiliation Department of Neurology, Lurie Comprehensive Cancer Center, Center for Genetic Medicine, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ML JYW. Performed the experiments: NH JW KF MV. Analyzed the data: ML AF JW JYW ZK WG MV. Wrote the paper: ML AF JW JYW ZK WG MV.