About the Authors

Wei Hou

Affiliation Department of Epidemiology and Health Policy Research, University of Florida, Gainesville, Florida, United States of America

John Stephen F. Yap

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

Song Wu

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

Tian Liu

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

James M. Cheverud

Affiliation Department of Anatomy and Neurobiology, Washington University Medical School, St. Louis, Missouri, United States of America

Rongling Wu

To whom correspondence should be addressed. E-mail: rwu@mail.ifas.ufl.edu

Affiliation Department of Statistics, University of Florida, Gainesville, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW. Performed the experiments: JC. Analyzed the data: WH SW TL. Wrote the paper: RW.