About the Authors

Sarah Knox

Contributed equally to this work with: Sarah Knox, Hong Ge

Affiliation The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America

Hong Ge

Contributed equally to this work with: Sarah Knox, Hong Ge

Affiliation The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America

Brian D. Dimitroff

Affiliations The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America, The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Yi Ren

Affiliations The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America, The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Katie A. Howe

Affiliations The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America, The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Andrew M. Arsham

Affiliation The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Mathew C. Easterday

Affiliations The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America, The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Thomas P. Neufeld

Affiliation The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Michael B. O'Connor

Affiliation The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Scott B. Selleck

To whom correspondence should be addressed. E-mail: selle011@umn.edu

Affiliations The Developmental Biology Center, Department of Pediatrics, The University of Minnesota, Minneapolis, Minnesota, United States of America, The Developmental Biology Center, Department of Genetics, Cell Biology and Development, The University of Minnesota, Minneapolis, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS MO MO SK AA TN. Performed the experiments: SK HG BD YR KH AA ME. Analyzed the data: SS MO MO SK HG BD YR ME TN. Wrote the paper: SS.