About the Authors

Kevin T. Williams

Affiliation The Liver Research Group and MRC Centre for Immune Regulation, Institute of Biomedical Research, The University of Birmingham, Birmingham, United Kingdom

Steven P. Young

Affiliation Department of Rheumatology, University of Birmingham Medical School, Birmingham, United Kingdom

Alison Negus

Affiliation The Liver Research Group and MRC Centre for Immune Regulation, Institute of Biomedical Research, The University of Birmingham, Birmingham, United Kingdom

Lawrence S. Young

Affiliation Cancer Research UK Institute for Cancer Studies, University of Birmingham Medical School, Birmingham, United Kingdom

David H. Adams

Affiliation The Liver Research Group and MRC Centre for Immune Regulation, Institute of Biomedical Research, The University of Birmingham, Birmingham, United Kingdom

Simon C. Afford

To whom correspondence should be addressed. E-mail: s.c.afford@bham.ac.uk

Affiliation The Liver Research Group and MRC Centre for Immune Regulation, Institute of Biomedical Research, The University of Birmingham, Birmingham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SA KW. Performed the experiments: KW SY AN. Analyzed the data: SA KW SY AN. Wrote the paper: SA. Other: Offered expert advice on interpretation of data and manuscript preparation: DA LY.