About the Authors

Elene R. Valdivia

Current address: Division of Biological Sciences, University of Missouri, Columbia, Missouri, United States of America

Affiliation Department of Biology and The Plant Physiology Graduate Program, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Yajun Wu

Current address: Department of Plants, Soils and Biometeorology, Utah State University, Logan, Utah, United States of America

Affiliation Department of Biology and The Plant Physiology Graduate Program, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Lian-Chao Li

Affiliation Proteomics and Mass Spectrometry Core Facility, The Huck Institutes of the Life Sciences, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Daniel J. Cosgrove

Affiliation Department of Biology and The Plant Physiology Graduate Program, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Andrew G. Stephenson

To whom correspondence should be addressed. E-mail: as4@psu.edu

Affiliation Department of Biology and The Plant Physiology Graduate Program, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AS EV LL DC. Performed the experiments: EV YW LL. Analyzed the data: AS EV. Contributed reagents/materials/analysis tools: DC. Wrote the paper: AS DC. Other: Assisted in manuscript preparation: L-CL DC EV.