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Fig 1.

Adapted from Shugart et al., 2020 [47], schematic showing how vegetation processes scale across space and time.

Grey boxes classify the biological and community scale. Yellow text describes leaf-based processes. Blue text describes endogenous processes. Red text describes exogenous disturbances and processes. The black dotted lines illustrate the current spatial and temporal scope of Earth System Models.

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Fig 2.

A rudimentary simulation of a forest from bare soil for 600 years, using the continuous patch Eq (2) along with a toy gap model using a fixed 50x50 grid of patches on an arbitrary scale, and for a patch disturbance rate of λ = 0.01yr−1.

Panel (a) shows how the age-distribution of patches changes through time for both the gap and statistical model. (b) shows the time-evolving spatial age map for the 50x50 grid.

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Fig 3.

Adapted from the work of Moore et al., 2018, 2020 [141, 142].

Best fits of Demographic Equilibrium Theory (DET), Eq (5), for μ0 (assuming zd and ϕ → 1/3) against observations of number density versus diameter. Panel (a) shows the DET fit to North American trees [141] observations of trunk diameter (diameter at breast height) from the USDA Forest Service FIA program [143]. Panel (b) presents DET fits to RAINFOR measurements of basal diameter (b) [144]. μ1 is the normalised metric of the turnover parameter (μ0 at d0 = 1 cm) describing the shape of the Weibull distribution.

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Fig 4.

Dynamic Global Vegetation Model (DGVM) trends through time.

Here models are categorized into three types: Average, Intermediate, and Complex. (1) Simple—plant size or age demographics are not included beyond the “average” case. (2) Intermediate—models include the representation of size and/or age within cohorts. (3) Complex—Individual Based models.

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