About the Authors

Kouki Shioya

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Charlotte Michaux

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Carsten Kuenne

Affiliation Institute of Medical Microbiology, Justus-Liebig-University, Giessen, Germany

Torsten Hain

Affiliation Institute of Medical Microbiology, Justus-Liebig-University, Giessen, Germany

Nicolas Verneuil

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Aurélie Budin-Verneuil

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Thomas Hartsch

Affiliation Genedata AG, Basel, Switzerland

Axel Hartke

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Jean-Christophe Giard

jean-christophe.giard@unicaen.fr

Affiliation Laboratoire de Microbiologie de l'Environnement, EA956-USC INRA 2017-IFR146 ICORE, University of Caen, Caen, France

Competing Interests

Thomas Hartsch is an employee of Genedata AG (Basel, Switzerland), and internal legal approval was granted to publish this study. It was a collaborative effort between University of Caen, University of Giessen, and Genedata company groups and is not related to any products manufactured or marketed. There are no patents to declare. The authors hereby confirm that studies presented do not alter adherence to all the PLoS ONE policies on sharing data and materials, as detailed in the PLoS ONE guide for authors.

Author Contributions

Conceived and designed the experiments: KS JCG TH AH. Performed the experiments: KS CM CK AB-V. Analyzed the data: KS CM NV JCG. Contributed reagents/materials/analysis tools: CK TH NV. Wrote the paper: KS AH JCG.