About the Authors

Hui-Yun Tseng

Affiliations Department of Life Science, Tunghai University, Taiwan, Department of Biology, National Museum of Natural Science, Taiwan

Chung-Ping Lin

wshuang@mail.nmns.edu.tw (WSH); treehops@thu.edu.tw (CPL)

Affiliations Department of Life Science, Tunghai University, Taiwan, Department of Life Science, National Taiwan Normal University, Taiwan

Jung-Ya Hsu

Affiliation Department of Biology, National Museum of Natural Science, Taiwan

David A. Pike

Affiliation School of Marine and Tropical Biology and Centre for Tropical Environmental and Sustainability Science, James Cook University, Townsville, Australia

Wen-San Huang

wshuang@mail.nmns.edu.tw (WSH); treehops@thu.edu.tw (CPL)

Affiliations Department of Biology, National Museum of Natural Science, Taiwan, Department of Life Sciences, National Chung Hsing University, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: WSH CPL DP HYT. Contributed reagents/materials/analysis tools: WSH CPL DP HYT. Wrote the paper: WSH CPL DP HYT. Designed research: WSH CPL HYT. Performed research: WSH JYH HYT.