About the Authors

Yoshihiro Matsuoka

matsuoka@fpu.ac.jp

Affiliation Fukui Prefectural University, Matsuoka, Eiheiji, Yoshida, Fukui, Japan

Shigeo Takumi

Affiliation Laboratory of Plant Genetics, Graduate School of Agricultural Science, Kobe University, Nada-ku, Kobe, Japan

Taihachi Kawahara

Affiliation Graduate School of Agriculture, Kyoto University, Muko, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YM ST TK. Performed the experiments: YM ST. Analyzed the data: YM ST. Contributed reagents/materials/analysis tools: YM TK. Wrote the paper: YM.