About the Authors

Martin Sikora

msikora@stanford.edu (MS); cdbustam@stanford.edu (CDB)

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Meredith L. Carpenter

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Andres Moreno-Estrada

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Brenna M. Henn

Current address: Department of Ecology and Evolution, Stony Brook University, Stony Brook, New York, United States of America

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Peter A. Underhill

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Federico Sánchez-Quinto

Affiliation Institute of Evolutionary Biology (CSIC-UPF), PRBB, Barcelona, Spain

Ilenia Zara

Affiliation CRS4 (Centre for Advanced Studies, Research and Development in Sardinia), Pula, Italy

Maristella Pitzalis

Affiliation Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy

Carlo Sidore

Affiliations Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America, Università degli Studi di Sassari, Dip. Scienze Biomediche, Sassari, Italy

Fabio Busonero

Affiliations Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Andrea Maschio

Affiliations Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Andrea Angius

Affiliations Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy, Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Chris Jones

Affiliation CRS4 (Centre for Advanced Studies, Research and Development in Sardinia), Pula, Italy

Javier Mendoza-Revilla

Affiliation Laboratorios de Investigación y Desarrollo, Facultad de Ciencias y Filosofía, Universidad Peruana Cayetano Heredia, Lima, Peru

Georgi Nekhrizov

Affiliation National Archaeological Institute with Museum, Sofia, Bulgaria

Diana Dimitrova

Affiliation National Archaeological Institute with Museum, Sofia, Bulgaria

Nikola Theodossiev

Affiliation Sofia University St. Kliment Ohridski, Sofia, Bulgaria

Timothy T. Harkins

Affiliation Life Technologies, Beverly, Massachusetts, United States of America

Andreas Keller

Affiliation University Hospital, Saarland University, Saarbrücken, Germany

Frank Maixner

Affiliation EURAC, Institute for mummies and the Iceman, Bolzano, Italy

Albert Zink

Affiliation EURAC, Institute for mummies and the Iceman, Bolzano, Italy

Goncalo Abecasis

Affiliation Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, United States of America

Serena Sanna

Affiliation Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy

Francesco Cucca

Affiliation Istituto di Ricerca Genetica e Biomedica (IRGB), CNR, Monserrato, Italy

Carlos D. Bustamante

msikora@stanford.edu (MS); cdbustam@stanford.edu (CDB)

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MS TTH AK AZ GA SS FC CDB BMH PAU. Performed the experiments: MLC IZ MP CS FB AM AA CJ. Analyzed the data: MS MLC AME FSQ IZ JMR. Contributed reagents/materials/analysis tools: FSQ IZ MP CS FB AM AA CJ GN DD NT FM. Wrote the paper: MS BMH PAU CDB.