About the Authors

Lina Fu

Contributed equally to this work with: Lina Fu, Yuqiong Hu

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft, Writing – review & editing

Affiliations National Laboratory of Biomacromolecules, CAS Center for Excellence in Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences, Beijing, China, State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Yuqiong Hu

Contributed equally to this work with: Lina Fu, Yuqiong Hu

Roles Data curation, Formal analysis, Investigation, Methodology, Writing – original draft

Affiliations Beijing Advanced Innovation Center for Genomics, College of Life Sciences, Peking University, Beijing, China, Biomedical Pioneering Innovation Center, Peking University, Beijing, China

Moshi Song

Roles Formal analysis, Writing – review & editing

Affiliations University of Chinese Academy of Sciences, Beijing, China, State Key Laboratory of Membrane Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, Institute for Stem Cell and Regeneration, Chinese Academy of Sciences, Beijing, China

Zunpeng Liu

Roles Data curation, Formal analysis, Visualization

Affiliations State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China

Weiqi Zhang

Roles Data curation, Formal analysis, Supervision, Writing – original draft

Affiliations National Laboratory of Biomacromolecules, CAS Center for Excellence in Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China, Institute for Stem Cell and Regeneration, Chinese Academy of Sciences, Beijing, China, Advanced Innovation Center for Human Brain Protection, National Clinical Research Center for Geriatric Disorders, Xuanwu Hospital Capital Medical University, Beijing, China

Fa-Xing Yu

Roles Writing – original draft

Affiliation Children's Hospital and Institutes of Biomedical Sciences, Fudan University, Shanghai, China

Jun Wu

Roles Writing – original draft, Writing – review & editing

Affiliation Department of Molecular Biology, University of Texas Southwestern Medical Center, Dallas, Texas, United States of America

Si Wang

Roles Methodology, Writing – review & editing

Affiliations National Laboratory of Biomacromolecules, CAS Center for Excellence in Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences, Beijing, China, Advanced Innovation Center for Human Brain Protection, National Clinical Research Center for Geriatric Disorders, Xuanwu Hospital Capital Medical University, Beijing, China

Juan Carlos Izpisua Belmonte

Roles Formal analysis, Methodology, Supervision

Affiliation Gene Expression Laboratory, Salk Institute for Biological Studies, La Jolla, California, United States of America

Piu Chan

Roles Formal analysis, Methodology, Supervision

Affiliation Advanced Innovation Center for Human Brain Protection, National Clinical Research Center for Geriatric Disorders, Xuanwu Hospital Capital Medical University, Beijing, China

Jing Qu

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Supervision, Writing – original draft, Writing – review & editing

qujing@ioz.ac.cn (JQ); tangfuchou@pku.edu.cn (FT); ghliu@ibp.ac.cn (GHL)

Affiliations State Key Laboratory of Stem Cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China, Institute for Stem Cell and Regeneration, Chinese Academy of Sciences, Beijing, China

Fuchou Tang

Roles Conceptualization, Formal analysis, Funding acquisition, Resources, Supervision, Validation, Writing – review & editing

qujing@ioz.ac.cn (JQ); tangfuchou@pku.edu.cn (FT); ghliu@ibp.ac.cn (GHL)

Affiliations Beijing Advanced Innovation Center for Genomics, College of Life Sciences, Peking University, Beijing, China, Biomedical Pioneering Innovation Center, Peking University, Beijing, China, Peking-Tsinghua Center for Life Sciences, Peking University, Beijing, China, Ministry of Education Key Laboratory of Cell Proliferation and Differentiation, Beijing, China

Guang-Hui Liu

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

qujing@ioz.ac.cn (JQ); tangfuchou@pku.edu.cn (FT); ghliu@ibp.ac.cn (GHL)

Affiliations National Laboratory of Biomacromolecules, CAS Center for Excellence in Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences, Beijing, China, University of Chinese Academy of Sciences, Beijing, China, State Key Laboratory of Membrane Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, Institute for Stem Cell and Regeneration, Chinese Academy of Sciences, Beijing, China, Advanced Innovation Center for Human Brain Protection, National Clinical Research Center for Geriatric Disorders, Xuanwu Hospital Capital Medical University, Beijing, China, Beijing Institute for Brain Disorders, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.