About the Authors

Simon K Cheng

Affiliation Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, and Department of Cell Biology, New York University School of Medicine, New York, New York, United States of America

Felix Olale

Affiliation Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, and Department of Cell Biology, New York University School of Medicine, New York, New York, United States of America

Ali H Brivanlou

Affiliation Laboratory of Molecular Vertebrate Embryology, The Rockefeller University, New York, New York, United States of America

Alexander F Schier

To whom correspondence should be addressed. E-mail: schier@saturn.med.nyu.edu

Affiliation Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, and Department of Cell Biology, New York University School of Medicine, New York, New York, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

SKC, AHB, and AFS conceived and designed the experiments. SKC and FO performed the experiments. SKC, FO, AHB, and AFS analyzed the data. SKC, FO, AHB, and AFS contributed reagents/materials/analysis tools. SKC and AFS wrote the paper.