About the Authors

Ming-Xian Chang

Contributed equally to this work with: Ming-Xian Chang, Jun Zou

Affiliations Scottish Fish Immunology Research Centre, University of Aberdeen, Aberdeen, United Kingdom, State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, China

Jun Zou

Contributed equally to this work with: Ming-Xian Chang, Jun Zou

Affiliation Scottish Fish Immunology Research Centre, University of Aberdeen, Aberdeen, United Kingdom

Pin Nie

Affiliation State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, China

Bei Huang

Affiliation College of Fisheries, Jimei University, Xiamen, China

Zhanglong Yu

Affiliation State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan, China

Bertrand Collet

Affiliation Marine Scotland Science Marine Laboratory, Aberdeen, United Kingdom

Chris J. Secombes

c.secombes@abdn.ac.uk

Affiliation Scottish Fish Immunology Research Centre, University of Aberdeen, Aberdeen, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JZ CJS. Performed the experiments: MXC JZ BC ZY. Analyzed the data: MXC JZ PN BH CJS. Wrote the paper: MXC JZ PN CJS.