About the Authors

Harriet Auty

Current address: Epidemiology Research Unit, Scottish Agricultural College, Inverness, United Kingdom

Affiliations Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom, Institute for Biodiversity, Animal Health and Comparative Medicine, College of Medicine, Veterinary Medicine and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Neil E. Anderson

Affiliation Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Kim Picozzi

Affiliation Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Tiziana Lembo

Affiliations Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom, Institute for Biodiversity, Animal Health and Comparative Medicine, College of Medicine, Veterinary Medicine and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Joseph Mubanga

Affiliation Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Richard Hoare

Affiliation Tanzania Wildlife Research Institute - Messerli Foundation Wildlife Veterinary Programme, Arusha, Tanzania

Robert D. Fyumagwa

Affiliation Tanzania Wildlife Research Institute - Serengeti Wildlife Research Centre, Arusha, Tanzania

Barbara Mable

Affiliation Institute for Biodiversity, Animal Health and Comparative Medicine, College of Medicine, Veterinary Medicine and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Louise Hamill

Affiliation Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Sarah Cleaveland

Affiliations Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom, Institute for Biodiversity, Animal Health and Comparative Medicine, College of Medicine, Veterinary Medicine and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Susan C. Welburn

sue.welburn@ed.ac.uk

Affiliation Centre for Infectious Diseases, College of Medicine and Veterinary Medicine, The University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HA NEA KP JM SC SCW. Performed the experiments: HA NEA KP TL JM RH RDF LH. Analyzed the data: HA NEA BM. Contributed reagents/materials/analysis tools: KP BM RH RDF SC SCW. Wrote the paper: HA NEA KP TL JM BM SC SCW.