About the Authors

Jennifer F. Dennis

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas Medical Center, Kansas City, Kansas, United States of America

Hiroshi Kurosaka

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Angelo Iulianella

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Neurobiology, Faculty of Medicine, Dalhousie University, Halifax, Nova Scotia, Canada

Jennifer Pace

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Nancy Thomas

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Sharon Beckham

Affiliation Stowers Institute for Medical Research, Kansas City, Missouri, United States of America

Trevor Williams

Affiliation Department of Cell Biology, Stem Cells, and Development, University of Colorado Denver, Aurora, Colorado, United States of America

Paul A. Trainor

pat@stowers.org

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas Medical Center, Kansas City, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFD HK PAT. Performed the experiments: JFD HK AI JP NT SB PAT. Analyzed the data: JFD HK AI JP TW PAT. Contributed reagents/materials/analysis tools: TW. Wrote the paper: JFD HK AI TW PAT.