About the Authors

Kun Wang

Contributed equally to this work with: Kun Wang, Stanley Langevin, Michael Kirby

Affiliations Department of Mathematics, Colorado State University, Fort Collins, CO, United States of America, Department of Mechanical Engineering & Materials Science, Yale University, New Haven, CT, United States of America

Stanley Langevin

Contributed equally to this work with: Kun Wang, Stanley Langevin, Michael Kirby

Affiliation Department of Microbiology, School of Medicine, University of Washington, Seattle, WA, United States of America

Corey S. O’Hern

Affiliations Department of Mechanical Engineering & Materials Science, Yale University, New Haven, CT, United States of America, Department of Applied Physics, Department of Physics, and Graduate Program in Computational Biology & Bioinformatics, Yale University, New Haven, CT, United States of America

Mark D. Shattuck

Affiliations Department of Mechanical Engineering & Materials Science, Yale University, New Haven, CT, United States of America, Department of Physics and Benjamin Levich Institute, The City College of the City University of New York, New York, NY, United States of America

Serenity Ogle

Affiliation Department of Biomedical Sciences, Colorado State University, Fort Collins, CO, United States of America

Adriana Forero

Affiliation Department of Microbiology, School of Medicine, University of Washington, Seattle, WA, United States of America

Juliet Morrison

Affiliation Department of Microbiology, School of Medicine, University of Washington, Seattle, WA, United States of America

Richard Slayden

Affiliation Department of Microbiology, Immunology and Pathology, Colorado State University, Fort Collins, CO, United States of America

Michael G. Katze

Affiliation Department of Microbiology, School of Medicine, University of Washington, Seattle, WA, United States of America

Michael Kirby

Contributed equally to this work with: Kun Wang, Stanley Langevin, Michael Kirby

Michael.Kirby@Colostate.edu

Affiliations Department of Mathematics, Colorado State University, Fort Collins, CO, United States of America, Department of Computer Science, Colorado State University, Fort Collins, CO, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KW SL CSO MDS RS MGK MK. Performed the experiments: KW. Analyzed the data: KW SL MK. Contributed reagents/materials/analysis tools: KW SL AF MK. Wrote the paper: KW SL MK SO. Contributed to Fig 8: AF JM.