About the Authors

Shiori Yabe

Current address: Agricultural Research Center, National Agriculture and Food Research Organization, Kannondai, Tsukuba, Ibaraki, Japan

Affiliation Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Science, The University of Tokyo, Bunkyo, Tokyo, Japan

Masanori Yamasaki

Affiliation Food Resources Education and Research Center, Graduate School of Agricultural Science, Kobe University, Kasai, Hyogo, Japan

Kaworu Ebana

Affiliation Genetic Resources Center, National Institute of Agrobiological Sciences, Tsukuba, Ibaraki, Japan

Takeshi Hayashi

Affiliation Agricultural Research Center, National Agriculture and Food Research Organization, Kannondai, Tsukuba, Ibaraki, Japan

Hiroyoshi Iwata

aiwata@mail.ecc.u-tokyo.ac.jp

Affiliation Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Science, The University of Tokyo, Bunkyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SY HI. Performed the experiments: SY HI. Analyzed the data: SY. Contributed reagents/materials/analysis tools: KE MY TH. Wrote the paper: SY KE MY TH HI.