About the Authors

Shigeru Sato

ssato@jichi.ac.jp

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Hiroshi Yajima

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Yasuhide Furuta

Affiliation Animal Resource Development Unit and Genetic Engineering Team, Division of Bio-function Dynamics Imaging, RIKEN Center for Life Science Technologies (CLST), Kobe, Hyogo, Japan

Keiko Ikeda

Affiliation Division of Biology, Hyogo College of Medicine, Nishinomiya, Hyogo, Japan

Kiyoshi Kawakami

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS KK. Performed the experiments: SS HY KI. Analyzed the data: SS. Wrote the paper: SS KK. Generated transgenic mice: YF.