About the Authors

Simon J. Girling

sgirling@rzss.org.uk

Affiliation Veterinary Department, Living Collection, Royal Zoological Society of Scotland, Edinburgh, United Kingdom

Roisin Campbell-Palmer

Affiliations Faculty of Arts and Sciences, Department of Environmental and Health Studies, Telemark University College, Bø i Telemark, Norway, Conservation Programmes, Royal Zoological Society of Scotland, Edinburgh, United Kingdom

Romain Pizzi

Affiliation Veterinary Department, Living Collection, Royal Zoological Society of Scotland, Edinburgh, United Kingdom

Mary A. Fraser

Affiliation School of Life, Sport and Social Science, Edinburgh Napier University, Sighthill Campus, Edinburgh, United Kingdom

Jonathan Cracknell

Affiliation Longleat Safari and Adventure Park, Longleat, Warminster, Wiltshire, United Kingdom

Jon Arnemo

Affiliations Department of Forestry and Wildlife Management, Faculty of Applied Ecology and Agricultural Sciences, Hedmark University College, Campus Evenstad, Koppang, Norway, Department of Wildlife, Fish and Environmental Studies, Faculty of Forest Sciences, Swedish University of Agricultural Sciences, Umeå, Sweden

Frank Rosell

Affiliation Conservation Programmes, Royal Zoological Society of Scotland, Edinburgh, United Kingdom

Competing Interests

Longleat Safari and Adventure Park provided support in the form of salaries for authors [JC], but did not have any additional role in the study design, data collection and analysis, decision to publish, or preparation of the manuscript. The specific roles of these authors are articulated in the ‘author contributions’ section. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SG RCP RP. Performed the experiments: SG RCP RP MF JC JA FR. Analyzed the data: SG RCP MF. Contributed reagents/materials/analysis tools: SG RCP RP JC JA FR. Wrote the paper: SG RCP MF.