About the Authors

Zhiguo E

Affiliation China National Rice Research Institute, 359 Tiyuchang Road, Hangzhou, 310006, China

Yuping Zhang

Affiliation China National Rice Research Institute, 359 Tiyuchang Road, Hangzhou, 310006, China

Tingting Li

Affiliation China National Rice Research Institute, 359 Tiyuchang Road, Hangzhou, 310006, China

Lei Wang

wanglei05@caas.cn (LW); zhaoheming@whu.edu.cn (HMZ)

Affiliation China National Rice Research Institute, 359 Tiyuchang Road, Hangzhou, 310006, China

Heming Zhao

wanglei05@caas.cn (LW); zhaoheming@whu.edu.cn (HMZ)

Affiliation Center for Genomics and Biotechnology, Fujian Agriculture and Forestry University, Fuzhou, 350002, Fujian Province, China

Competing Interests

Co-author Heming Zhao is employed by Life Science and Technology Center, China National Seed Group Co., Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ZGE LW HMZ. Performed the experiments: ZGE YPZ TTL. Analyzed the data: ZGE TTL. Contributed reagents/materials/analysis tools: ZGE TTL HMZ. Wrote the paper: ZGE HMZ.