About the Authors

Tianqi Jia

Affiliation College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou, Fujian, P. R. China

Danfeng Wei

Affiliation College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou, Fujian, P. R. China

Shan Meng

Affiliation College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou, Fujian, P. R. China

Andrew C. Allan

Affiliation The New Zealand Institute for Plant & Food Research Limited (Plant & Food Research), Mt Albert, Auckland, New Zealand

Lihui Zeng

lhzeng@hotmail.com

Affiliation College of Horticulture, Fujian Agriculture and Forestry University, Fuzhou, Fujian, P. R. China

Competing Interests

Andrew C. Allan is an employee of The New Zealand Institute for Plant & Food Research Limited and received salary to fund this research. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LZ. Performed the experiments: TJ DW SM. Analyzed the data: TJ DW SM LZ. Contributed reagents/materials/analysis tools: TJ DW SM. Wrote the paper: LZ ACA.