About the Authors

Katherine L. Irvine

Contributed equally to this work with: Katherine L. Irvine, Monique Gangloff

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Monique Gangloff

Contributed equally to this work with: Katherine L. Irvine, Monique Gangloff

Affiliation Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Catherine M. Walsh

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

David R. Spring

Affiliation Department of Chemistry, University of Cambridge, Cambridge, United Kingdom

Nicholas J. Gay

Affiliation Department of Biochemistry, University of Cambridge, Cambridge, United Kingdom

Clare E. Bryant

ceb27@cam.ac.uk

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors would like to confirm that Nicholas J. Gay is a PLOS ONE Editorial Board member, and that this does not alter their adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: CEB. Performed the experiments: KLI MG. Analyzed the data: KLI MG NJG. Contributed reagents/materials/analysis tools: CMW. Wrote the paper: KLI MG CMW DRS NJG CEB. Generated structural images: DRS.