About the Authors

James F. Castellano

Contributed equally to this work with: James F. Castellano, Bonnie R. Fletcher

Affiliations Laboratory of Experimental Gerontology, National Institute on Aging, Baltimore, Maryland, United States of America, Graduate Program in Neuroscience, Mount Sinai School of Medicine, New York, New York, United States of America

Bonnie R. Fletcher

Contributed equally to this work with: James F. Castellano, Bonnie R. Fletcher

Affiliation Laboratory of Experimental Gerontology, National Institute on Aging, Baltimore, Maryland, United States of America

Bennett Kelley-Bell

Affiliation Laboratory of Experimental Gerontology, National Institute on Aging, Baltimore, Maryland, United States of America

David H. Kim

Affiliation Laboratory of Experimental Gerontology, National Institute on Aging, Baltimore, Maryland, United States of America

Michela Gallagher

Affiliation Department of Psychological and Brain Sciences, Johns Hopkins University, Baltimore, Maryland, United States of America

Peter R. Rapp

rappp@mail.nih.gov

Affiliation Laboratory of Experimental Gerontology, National Institute on Aging, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JFC BF PR MG. Performed the experiments: JFC BF DK BK. Analyzed the data: JFC BF PR. Contributed reagents/materials/analysis tools: JFC PR MG. Wrote the paper: JFC BF PR.