About the Authors

Wenting Wu

Contributed equally to this work with: Wenting Wu, Huan Li

Affiliations State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China, Beyster Center for Genomics of Psychiatric Diseases, Department of Psychiatry, University of California San Diego, La Jolla, California, United States of America

Huan Li

Contributed equally to this work with: Wenting Wu, Huan Li

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Huibo Wang

Affiliation Division of Hematology and Oncology, Comprehensive Cancer Center, University of Alabama at Birmingham, Birmingham, Alabama, United States of America

Xueying Zhao

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Zhiqiang Gao

Affiliation Department of Respiratory Disease, Shanghai Chest Hospital, Shanghai Jiaotong University, Shanghai, China

Rong Qiao

Affiliation Department of Respiratory Disease, Shanghai Chest Hospital, Shanghai Jiaotong University, Shanghai, China

Wei Zhang

Affiliation Department of Respiratory Disease, Shanghai Chest Hospital, Shanghai Jiaotong University, Shanghai, China

Ji Qian

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Jiucun Wang

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Hongyan Chen

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Qingyi Wei

Affiliation Department of Epidemiology, University of Texas M.D. Anderson Cancer Center, Houston, Texas, United States of America

Baohui Han

drlu@fudan.edu.cn (DL); xkyyhan@gmail.com(BH)

Affiliation Department of Respiratory Disease, Shanghai Chest Hospital, Shanghai Jiaotong University, Shanghai, China

Daru Lu

drlu@fudan.edu.cn (DL); xkyyhan@gmail.com(BH)

Affiliation State Key Laboratory of Genetic Engineering, Center for Fudan-VARI Genetic Epidemiology and MOE Key Laboratory of Contemporary Anthropology, Institute of Genetics, School of Life Sciences, Fudan University, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WTW HL DRL BHH QYW. Performed the experiments: WTW HL HBW ZQG RQ WZ. Analyzed the data: WTW HL XYZ QYW. Contributed reagents/materials/analysis tools: WTW HL JQ JCW HYC. Wrote the paper: DRL BHH WTW HL.