About the Authors

Rui Li

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation Departments of Medicine, Human Genetics, Epidemiology, and Biostatistics, Lady Davis Institute, Jewish General Hospital, McGill University, Montreal, Quebec, Canada

Felix F. Brockschmidt

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliations Institute of Human Genetics, University of Bonn, Bonn, Germany, Department of Genomics, Life and Brain Center, University of Bonn, Bonn, Germany

Amy K. Kiefer

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation 23andMe, Mountain View, California, United States of America

Hreinn Stefansson

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation deCODE genetics, Reykjavík, Iceland

Dale R. Nyholt

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Kijoung Song

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation Genetics Division, GlaxoSmithKline, King of Prussia, Pennsylvania, United States of America

Sita H. Vermeulen

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliations Department of Epidemiology, Biostatistics, and HTA, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Department of Genetics, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Stavroula Kanoni

Contributed equally to this work with: Rui Li, Felix F. Brockschmidt, Amy K. Kiefer, Hreinn Stefansson, Dale R. Nyholt, Kijoung Song, Sita H. Vermeulen, Stavroula Kanoni

Affiliation Genetics of Complex Traits in Humans, Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Daniel Glass

Affiliation Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Sarah E. Medland

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Maria Dimitriou

Affiliation Department of Dietetics-Nutrition, Harokopio University, Athens, Greece

Dawn Waterworth

Affiliation Genetics Division, GlaxoSmithKline, King of Prussia, Pennsylvania, United States of America

Joyce Y. Tung

Affiliation 23andMe, Mountain View, California, United States of America

Frank Geller

Affiliation Department of Epidemiology Research, Statens Serum Institute, Copenhagen, Denmark

Stefanie Heilmann

Affiliations Institute of Human Genetics, University of Bonn, Bonn, Germany, Department of Genomics, Life and Brain Center, University of Bonn, Bonn, Germany

Axel M. Hillmer

Affiliation Genome Technology and Biology, Genome Institute of Singapore, Singapore, Singapore

Veronique Bataille

Affiliation Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Sibylle Eigelshoven

Affiliation Department of Dermatology, University of Düsseldorf, Düsseldorf, Germany

Sandra Hanneken

Affiliation Department of Dermatology, University of Düsseldorf, Düsseldorf, Germany

Susanne Moebus

Affiliation Institute for Medical Informatics, Biometry, and Epidemiology, University Clinic Essen, University Duisburg-Essen, Essen, Germany

Christine Herold

Affiliation German Center for Neurodegenerative Diseases (DZNE), Bonn, Germany

Martin den Heijer

Affiliation Department of Endocrinology, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands

Grant W. Montgomery

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Panos Deloukas

Affiliation Genetics of Complex Traits in Humans, Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Nicholas Eriksson

Affiliation 23andMe, Mountain View, California, United States of America

Andrew C. Heath

Affiliation Washington University Medical School, St. Louis, Missouri, United States of America

Tim Becker

Affiliations German Center for Neurodegenerative Diseases (DZNE), Bonn, Germany, Institute for Medical Biometry, Informatics, and Epidemiology, University of Bonn, Bonn, Germany

Patrick Sulem

Affiliation deCODE genetics, Reykjavík, Iceland

Massimo Mangino

Affiliation Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Peter Vollenweider

Affiliation Department of Internal Medicine, CHUV University Hospital, Lausanne, Switzerland

Tim D. Spector

Affiliation Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

George Dedoussis

These authors also contributed equally to this work.

Affiliation Department of Dietetics-Nutrition, Harokopio University, Athens, Greece

Nicholas G. Martin

These authors also contributed equally to this work.

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Lambertus A. Kiemeney

These authors also contributed equally to this work.

Affiliations Department of Epidemiology, Biostatistics, and HTA, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Department of Urology, Radboud University Nijmegen Medical Centre, Nijmegen, The Netherlands, Comprehensive Cancer Centre of the Netherlands (IKNL), Nijmegen, The Netherlands

Vincent Mooser

These authors also contributed equally to this work.

Affiliation Genetics Division, GlaxoSmithKline, King of Prussia, Pennsylvania, United States of America

Kari Stefansson

These authors also contributed equally to this work.

Affiliation deCODE genetics, Reykjavík, Iceland

David A. Hinds

These authors also contributed equally to this work.

Affiliation 23andMe, Mountain View, California, United States of America

Markus M. Nöthen

These authors also contributed equally to this work.

Affiliations Institute of Human Genetics, University of Bonn, Bonn, Germany, Department of Genomics, Life and Brain Center, University of Bonn, Bonn, Germany, German Center for Neurodegenerative Diseases (DZNE), Bonn, Germany

J. Brent Richards

brent.richards@mcgill.ca

These authors also contributed equally to this work.

Affiliations Departments of Medicine, Human Genetics, Epidemiology, and Biostatistics, Lady Davis Institute, Jewish General Hospital, McGill University, Montreal, Quebec, Canada, Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FF Brockschmidt, AK Kiefer, H Stefansson, DR Nyholt, D Waterworth, JY Tung, F Geller, AM Hillmer, V Bataille, GW Montgomery, N Eriksson, AC Heath, P Sulem, TD Spector, NG Martin, LA Kiemeney, V Moorser, K Stefansson, DA Hinds, MM Nöthen, JB Richards. Performed the experiments: FF Brockschmidt, H Stefansson, D Glass, S Heilmann, AM Hillmer, V Bataille, AC Heath, P Sulem, LA Kiemeney, V Mooser. Analyzed the data: R Li, FF Brockschmidt, H Stefansson, DR Nyholt, K Song, SH Vermeulen, S Kanoni, D Glass, SE Medland, D Waterworth, F Geller, S Heilmann, AM Hillmer, C Herold, M den Heijer, T Becker, P Sulem, M Mangino, P Vollenweider, LA Kiemeney, DA Hinds, JB Rrichards. Contributed reagents/materials/analysis tools: FF Brockschmidt, DR Nyholt, SE Medland, S Heilmann, AM Hillmer, S Eigelshoven, S Hanneken, S Moebus, GW Montgomery, P Deloukas, N Eriksson, AC Heath, TD Spector, G Dedoussis, NG Martin, K Stefansson, MM Nöthen. Wrote the paper: R Li, FF Brockschmidt, DR Nyholt, S Heilmann, AM Hillmer, K Stefansson, DA Hinds, MM Nöthen, JB Richards. Survey development: AK Kiefer, JY Tung. Phenotype collection/data entry: M Dimitriou. Recruitment and phenotyping of cases: S Eigelshoven, S Hanneken. Study coordinator: S Moebus.