About the Authors

Amir N. Saleem

Contributed equally to this work with: Amir N. Saleem, Yen-Hui Chen, Hwa Jin Baek

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Department of Internal and Preventive Medicine, College of Veterinary Medicine, University of Mosul, Mosul, Iraq

Yen-Hui Chen

Contributed equally to this work with: Amir N. Saleem, Yen-Hui Chen, Hwa Jin Baek

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Taiwan Mouse Clinic–National Phenotyping Center, National Research Program for Genomic Medicine, National Science Council, Taipei, Taiwan

Hwa Jin Baek

Contributed equally to this work with: Amir N. Saleem, Yen-Hui Chen, Hwa Jin Baek

Affiliation Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Ya-Wen Hsiao

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Taiwan Mouse Clinic–National Phenotyping Center, National Research Program for Genomic Medicine, National Science Council, Taipei, Taiwan

Hong-Wen Huang

Affiliation Taiwan Mouse Clinic–National Phenotyping Center, National Research Program for Genomic Medicine, National Science Council, Taipei, Taiwan

Hsiao-Jung Kao

Affiliation Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan

Kai-Ming Liu

Affiliation Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan

Li-Fen Shen

Affiliation Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan

I-wen Song

Affiliation Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan

Chen-Pei D. Tu

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Department of Biochemistry and Molecular Biology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Jer-Yuarn Wu

Affiliation Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan

Tateki Kikuchi

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Taiwan Mouse Clinic–National Phenotyping Center, National Research Program for Genomic Medicine, National Science Council, Taipei, Taiwan

Monica J. Justice

Affiliation Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Jeffrey J. Y. Yen

chen0010@ibms.sinica.edu.tw (Y-TC); bmjyen@ibms.sinica.edu.tw (JJYY)

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Taiwan Mouse Clinic–National Phenotyping Center, National Research Program for Genomic Medicine, National Science Council, Taipei, Taiwan

Yuan-Tsong Chen

chen0010@ibms.sinica.edu.tw (Y-TC); bmjyen@ibms.sinica.edu.tw (JJYY)

Affiliations Institute of Biomedical Sciences, Academia Sinica, Nankang, Taiwan, Department of Pediatrics, Duke University Medical Center, Durham, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ANS YTC. Performed the experiments: ANS YHC HJB YWH HWH KML LFS IwS MJJ. Analyzed the data: ANS HJK KML CPDT JYW TK JJYY. Contributed reagents/materials/analysis tools: JYW MJJ JJYY YTC. Wrote the paper: ANS YTC. Interpreted Histopathological and IHC findings: YHC.