About the Authors

Gordon T. Brown

Affiliation Pathology, Division of Applied Medicine, School of Medicine and Dentistry, University of Aberdeen, Aberdeen, United Kingdom

Beatriz Gimenez Cash

Affiliation Vertebrate Antibodies, Zoology Building, Tillydrone Avenue, Aberdeen, United Kingdom

Daniela Blihoghe

Affiliation George S. Wise Faculty of Life Sciences, Department of Zoology, Tel Aviv University, Tel Aviv, Israel

Petronella Johansson

Affiliation The Scottish Fish Immunology Research Centre, School of Biological Sciences, University of Aberdeen, Aberdeen, United Kingdom

Ayham Alnabulsi

Affiliation Vertebrate Antibodies, Zoology Building, Tillydrone Avenue, Aberdeen, United Kingdom

Graeme I. Murray

g.i.murray@abdn.ac.uk

Affiliation Pathology, Division of Applied Medicine, School of Medicine and Dentistry, University of Aberdeen, Aberdeen, United Kingdom

Competing Interests

Graeme Murray is a scientific advisor to Vertebrate Antibodies, which contributed to this research. No financial remuneration has been or is received for this position nor does he hold any shares in this company. Beatriz Cash and Ayham Alnabulsi are employees of Vertebrate Antibodies. This does not alter the authors' adherence to PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GTB BGC AA GIM. Performed the experiments: GTB BGC DB PJ AA GIM. Analyzed the data: GTB BGC AA GIM. Contributed reagents/materials/analysis tools: GTB BGC DB PJ AA GIM. Wrote the paper: GTB BGC AA GIM.