About the Authors

Wen-Jie Zhang

Contributed equally to this work with: Wen-Jie Zhang, Yuan Niu, Su-Hong Bu, Meng Li

Affiliations Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China, Institute of Crop Research, Ningxia Academy of Agriculture and Forestry Sciences, Yinchuan, Ningxia, China

Yuan Niu

Contributed equally to this work with: Wen-Jie Zhang, Yuan Niu, Su-Hong Bu, Meng Li

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Su-Hong Bu

Contributed equally to this work with: Wen-Jie Zhang, Yuan Niu, Su-Hong Bu, Meng Li

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Meng Li

Contributed equally to this work with: Wen-Jie Zhang, Yuan Niu, Su-Hong Bu, Meng Li

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Jian-Ying Feng

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Jin Zhang

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Sheng-Xian Yang

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Medrine Mmayi Odinga

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Shi-Ping Wei

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Xiao-Feng Liu

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Yuan-Ming Zhang

soyzhang@njau.edu.cn

Affiliation Section on Statistical Genomics, State Key Laboratory of Crop Genetics and Germplasm Enhancement, Department of Crop Genetics and Breeding, Nanjing Agricultural University, Nanjing, Jiangsu, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YMZ. Performed the experiments: WJZ YN SPW XFL SXY YMZ. Analyzed the data: YN SHB ML WJZ JYF JZ YMZ. Contributed reagents/materials/analysis tools: SHB ML. Wrote the paper: YMZ YN MMO.