About the Authors

Alexander H. J. Wittenberg

Contributed equally to this work with: Alexander H. J. Wittenberg, Theo A. J. van der Lee, Sarrah Ben M'Barek

Current address: Keygene N.V., Wageningen, The Netherlands

Affiliations Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands, Graduate School Experimental Plant Sciences, Laboratory of Plant Breeding, Department of Plant Sciences, Wageningen University and Research Centre, Wageningen, The Netherlands

Theo A. J. van der Lee

Contributed equally to this work with: Alexander H. J. Wittenberg, Theo A. J. van der Lee, Sarrah Ben M'Barek

Affiliation Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands

Sarrah Ben M'Barek

Contributed equally to this work with: Alexander H. J. Wittenberg, Theo A. J. van der Lee, Sarrah Ben M'Barek

Affiliations Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands, Graduate School Experimental Plant Sciences, Laboratory of Phytopathology, Department of Plant Sciences, Wageningen University and Research Centre, Wageningen, The Netherlands

Sarah B. Ware

Affiliations Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands, Graduate School Experimental Plant Sciences, Laboratory of Phytopathology, Department of Plant Sciences, Wageningen University and Research Centre, Wageningen, The Netherlands

Stephen B. Goodwin

Affiliation United States Department of Agriculture (USDA)-Agricultural Research Service (ARS), Crop Production and Pest Control Research Unit, and Department of Botany and Plant Pathology, Purdue University, West Lafayette, Indiana, United States of America

Andrzej Kilian

Affiliation Diversity Arrays P/L, Yarralumla, Canberra, Australian Capital Territory, Australia

Richard G. F. Visser

Affiliations Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands, Graduate School Experimental Plant Sciences, Laboratory of Plant Breeding, Department of Plant Sciences, Wageningen University and Research Centre, Wageningen, The Netherlands

Gert H. J. Kema

gert.kema@wur.nl

Affiliation Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands

Henk J. Schouten

Affiliation Plant Research International B.V., Wageningen University and Research Centre, Wageningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AHJW TAJVdL GHJK HJS. Performed the experiments: AHJW TAJVdL SBM SBW AK. Analyzed the data: AHJW TAJVdL SBM GHJK HJS. Contributed reagents/materials/analysis tools: AK. Wrote the paper: AHJW TAJVdL SBG RGFV GHJK HJS.