About the Authors

Chen Chu

Contributed equally to this work with: Chen Chu, Guangyong Zheng

Affiliations State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China, Graduate University of the Chinese Academy of Sciences, 200031, Shanghai, China

Guangyong Zheng

Contributed equally to this work with: Chen Chu, Guangyong Zheng

Affiliation CAS-MPG Partner Institute for Computational Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Shuanggang Hu

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Jinsong Zhang

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Shengsong Xie

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Wubin Ma

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Minjie Ni

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Chunhua Tang

Affiliations State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China, Graduate University of the Chinese Academy of Sciences, 200031, Shanghai, China

Lu Zhou

Affiliations Key Lab of Systems Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China, Graduate University of the Chinese Academy of Sciences, 200031, Shanghai, China

Yuchuan Zhou

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Mofang Liu

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Yixue Li

yxli@sibs.ac.cn (YXL); ylzhang@sibcb.ac.cn (YLZ)

Affiliation Key Lab of Systems Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Yonglian Zhang

yxli@sibs.ac.cn (YXL); ylzhang@sibcb.ac.cn (YLZ)

Affiliation State Key Laboratory of Molecular Biology, Shanghai Key Laboratory of Molecular Andrology, Institute of Biochemistry and Cell Biology, Shanghai Institutes for Biological Sciences, Chinese Academy of Sciences, 200031, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CC GZ SH JZ Y. Zhang. Performed the experiments: SH JZ CC SX WM MN CT. Analyzed the data: GZ LZ. Contributed reagents/materials/analysis tools: CC GZ SH JZ SX WM MN CT LZ Y. Zhou. Wrote the paper: CC GZ Y. Zhou ML YL Y. Zhang.