About the Authors

Kristi A. Koelsch

Kristi-Koelsch@omrf.org (KAK); jdwren@gmail.com (JDW)

Affiliation Oklahoma Medical Research Foundation, Arthritis and Clinical Immunology Program, Oklahoma City, Oklahoma, United States of America

YuJing Wang

Affiliation Oklahoma Medical Research Foundation, Arthritis and Clinical Immunology Program, Oklahoma City, Oklahoma, United States of America

Jacen S. Maier-Moore

Affiliations Oklahoma Medical Research Foundation, Arthritis and Clinical Immunology Program, Oklahoma City, Oklahoma, United States of America, University of Oklahoma Health Sciences Center, Department of Medicine, Oklahoma City, Oklahoma, United States of America

Amr H. Sawalha

Affiliations Oklahoma Medical Research Foundation, Arthritis and Clinical Immunology Program, Oklahoma City, Oklahoma, United States of America, Division of Rheumatology, Department of Internal Medicine, University of Michigan, Ann Arbor, Michigan, United States of America

Jonathan D. Wren

Kristi-Koelsch@omrf.org (KAK); jdwren@gmail.com (JDW)

Affiliations Oklahoma Medical Research Foundation, Arthritis and Clinical Immunology Program, Oklahoma City, Oklahoma, United States of America, University of Oklahoma Health Sciences Center, Department of Biochemistry and Molecular Biology, Oklahoma City, Oklahoma, United States of America

Competing Interests

AHS, M.D. is a PLOS ONE Editorial Board Member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Supervised experiments and data analysis: AHS JDW. Conceived and designed the experiments: KAK YW JSM. Performed the experiments: KAK YW JSM. Analyzed the data: KAK YW JSM. Contributed reagents/materials/analysis tools: JSM AHS JDW. Wrote the paper: KAK JSM.