About the Authors

Bahram Namjou

Bahram.namjou@cchmc.org

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America

Yizhao Ni

Affiliation Division of Biomedical Informatics, Cincinnati Children's Hospital Medical Center, Cincinnati, OH, 45229, United States of America

Isaac T. W. Harley

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America, Division of Rheumatology and Department of Internal Medicine, University of Colorado Denver, Aurora, CO, 80045, United States of America

Iouri Chepelev

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America

Beth Cobb

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America

Leah C. Kottyan

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America

Patrick M. Gaffney

Affiliation Arthritis and Clinical Immunology, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, 73104, United States of America

Joel M. Guthridge

Affiliation Arthritis and Clinical Immunology, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, 73104, United States of America

Kenneth Kaufman

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America

John B. Harley

Affiliations Center for Autoimmune Genomics and Etiology, Cincinnati Children's Hospital Medical Center (CCHMC), 3333 Burnet Avenue, Cincinnati, OH, 45229, United States of America, University of Cincinnati, College of Medicine, Cincinnati, OH, United States of America, Department of Veteran Affairs Medical Center, Cincinnati, OH, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BN IH. Performed the experiments: BN YN. Analyzed the data: BN YN. Contributed reagents/materials/analysis tools: JBH KK BN JG PG IC BC LK. Wrote the paper: BN BC IH JBH.