About the Authors

Toshihiro Imada

Affiliation Department of Ophthalmology, Keio University School of Medicine, Tokyo, Japan

Shigeru Nakamura

vdtwork@gmail.com

Affiliation Department of Ophthalmology, Keio University School of Medicine, Tokyo, Japan

Naoki Kitamura

Affiliation Department of Veterinary Physiology, Faculty of Agriculture, Tottori University, Tottori, Japan

Izumi Shibuya

Affiliation Department of Veterinary Physiology, Faculty of Agriculture, Tottori University, Tottori, Japan

Kazuo Tsubota

Affiliation Department of Ophthalmology, Keio University School of Medicine, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI SN KT. Performed the experiments: TI. Analyzed the data: TI SN. Wrote the paper: TI SN NK IS KT.