About the Authors

Ladd M. Irvine

ladd.irvine@oregonstate.edu

Affiliation Marine Mammal Institute, Department of Fisheries and Wildlife, and Coastal Oregon Marine Experiment Station, Oregon State University, Hatfield Marine Science Center, Newport, Oregon, United States of America

Bruce R. Mate

Affiliation Marine Mammal Institute, Department of Fisheries and Wildlife, and Coastal Oregon Marine Experiment Station, Oregon State University, Hatfield Marine Science Center, Newport, Oregon, United States of America

Martha H. Winsor

Affiliation Marine Mammal Institute, Department of Fisheries and Wildlife, and Coastal Oregon Marine Experiment Station, Oregon State University, Hatfield Marine Science Center, Newport, Oregon, United States of America

Daniel M. Palacios

Current address: Marine Mammal Institute, Department of Fisheries and Wildlife, and Coastal Oregon Marine Experiment Station, Oregon State University, Hatfield Marine Science Center, Newport, Oregon, United States of America

Affiliations Cooperative Institute for Marine Ecosystems and Climate, Institute of Marine Sciences, Division of Physical and Biological Sciences, University of California Santa Cruz, Santa Cruz, California, United States of America, NOAA/NMFS/SWFSC/Environmental Research Division, Pacific Grove, California, United States of America

Steven J. Bograd

Affiliation NOAA/NMFS/SWFSC/Environmental Research Division, Pacific Grove, California, United States of America

Daniel P. Costa

Affiliation Ecology and Evolutionary Biology, Long Marine Laboratory, University of California Santa Cruz, Santa Cruz, California, United States of America

Helen Bailey

Affiliation Chesapeake Biological Laboratory, University of Maryland Center for Environmental Science, Solomons, Maryland, United States of America

Competing Interests

Steven Bograd is currently an editor for this journal and is an author on this paper. This does not alter the authors’ adherence to PLOS ONE Editorial policies and criteria.

Author Contributions

Conceived and designed the experiments: LI BM. Performed the experiments: LI BM MW. Analyzed the data: LI HB DP. Contributed reagents/materials/analysis tools: HB. Wrote the paper: LI BM MW DP SB DC HB.