About the Authors

Haley D. O’Brien

ho128009@ohio.edu

Affiliation Ohio University, Department of Biological Sciences, Graduate Program in Ecology and Evolutionary Biology, Athens, Ohio, United States of America

Susan H. Williams

Affiliation Ohio University Heritage College of Osteopathic Medicine, Department of Biomedical Sciences, Athens, Ohio, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SHW HDO. Performed the experiments: SHW HDO. Analyzed the data: HDO. Contributed reagents/materials/analysis tools: HDO. Wrote the paper: HDO.