About the Authors

W. David Hill

Contributed equally to this work with: W. David Hill, Christiaan de Leeuw

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Christiaan de Leeuw

Contributed equally to this work with: W. David Hill, Christiaan de Leeuw

Affiliations Center for Neurogenomics and Cognitive Research, Neuroscience Campus Amsterdam, Complex Trait Genetics, VU University Amsterdam, Amsterdam, The Netherlands, Institute for Computing and Information Sciences, Radboud University Nijmegen, Nijmegen, The Netherlands

Gail Davies

Affiliation Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

David Cherry McLachlan Liewald

Affiliation Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

Anthony Payton

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

Leone C. A. Craig

Affiliation Public Health Nutrition Group, Institute of Applied Health Sciences, University of Aberdeen, Aberdeen, United Kingdom

Lawrence J. Whalley

Affiliation Institute of Applied Health Sciences, University of Aberdeen, Aberdeen, United Kingdom

Mike Horan

Affiliation Centre for Clinical and Cognitive Neurosciences, Institute of Brain, Behaviour and Mental Health, University of Manchester, Manchester, United Kingdom

William Ollier

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

John M. Starr

Affiliation Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom

Neil Pendleton

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

Danielle Posthuma

Affiliations Center for Neurogenomics and Cognitive Research, Neuroscience Campus Amsterdam, Complex Trait Genetics, VU University Amsterdam, Amsterdam, The Netherlands, Department of Clinical Genetics, VU University Medical Center, Amsterdam, the Netherlands, Department of Child and Adolescent Psychiatry, Erasmus University Rotterdam, Sophia Child Hospital, Rotterdam, The Netherlands

Timothy C. Bates

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Ian J. Deary

iand@staffmail.ed.ac.uk

Affiliations Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, United Kingdom, Department of Psychology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WDH TCB. Performed the experiments: WDH CdeL. Analyzed the data: WDH CdeL GD. Contributed reagents/materials/analysis tools: DCML LCAC AP MH WO NP DP IJD GD. Wrote the paper: WDH TCB IJD JMS LJW DP.