About the Authors

Chad Smithson

Affiliation Department of Biochemistry and Microbiology, University of Victoria, Victoria, British Columbia, Canada

Alex Purdy

Affiliation Department of Biochemistry and Microbiology, University of Victoria, Victoria, British Columbia, Canada

Adrian J. Verster

Affiliation Department of Molecular Genetics, University of Toronto, Toronto, Ontario, Canada

Chris Upton

cupton@uvic.ca

Affiliation Department of Biochemistry and Microbiology, University of Victoria, Victoria, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CU AJV. Performed the experiments: CS AJV. Analyzed the data: CU CS AJV. Contributed reagents/materials/analysis tools: AP. Wrote the paper: CU CS.