About the Authors

Hideaki Abe

Affiliations Wildlife Research Center, Kyoto University, Kyoto, Japan, Department of Anatomy, University of Otago, Dunedin, New Zealand

Kenji Nagao

Affiliation Animal Husbandry Research Division, Aichi Agricultural Research Center, Aichi, Japan

Miho Inoue-Murayama

mmurayama@wrc.kyoto-u.ac.jp

Affiliation Wildlife Research Center, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HA MI-M. Performed the experiments: HA KN. Analyzed the data: HA. Contributed reagents/materials/analysis tools: KN. Wrote the paper: HA. Conceived and designed molecular experiments: HA. Co-conceived all behavioral and molecular experiments: MI-M. Managed the domestic chicks used in behavioral experiments: KN. Performed molecular experiments: HA. Carried out behavioral experiments and qPCR: KN. Gave final approval of the version to be published: MI-M. Read and approved the final manuscript: MI-M KN HA.