About the Authors

Shang-Yin Vanson Liu

Affiliations Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan, Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, United States of America

Chia-Ling Carynn Chan

Affiliation Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan

Oceana Lin

Affiliation The Society of Wilderness, Taipei, Taiwan

Chieh-Shen Hu

Affiliation The Society of Wilderness, Taipei, Taiwan

Chaolun Allen Chen

cac@gate.sinica.edu.tw

Affiliations Biodiversity Research Center, Academia Sinica, Nangang, Taipei, Taiwan, Taiwan International Graduate Program-Biodiversity, Academia Sinica, Nangang, Taipei, Taiwan, Institute of Oceanography, National Taiwan University, Taipei, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CAC. Performed the experiments: CCC. Analyzed the data: CCC SVL. Contributed reagents/materials/analysis tools: OL CSH. Wrote the paper: SVL CCC CAC.