About the Authors

Hiroko Sano

sano_hiroko@kurume-u.ac.jp

Current address: Department of Molecular Genetics, Institute of Life Sciences, Kurume University, Kurume, Fukuoka, Japan

Affiliation HHMI and Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, Department of Cell Biology, New York University Medical Center, New York, New York, United States of America

Prabhat S. Kunwar

Contributed equally to this work with: Prabhat S. Kunwar, Andrew D. Renault

Current address: Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Affiliation HHMI and Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, Department of Cell Biology, New York University Medical Center, New York, New York, United States of America

Andrew D. Renault

Contributed equally to this work with: Prabhat S. Kunwar, Andrew D. Renault

Current address: Max Plank Institute for Developmental Biology, Tübingen, Germany

Affiliation HHMI and Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, Department of Cell Biology, New York University Medical Center, New York, New York, United States of America

Vitor Barbosa

Current address: Instituto Gulbenkian de Ciência, Oeiras, Portugal

Affiliation HHMI and Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, Department of Cell Biology, New York University Medical Center, New York, New York, United States of America

Ivan B. N. Clark

Affiliation Center for Systems Biology at Edinburgh, Edinburgh, United Kingdom

Shuji Ishihara

Affiliations Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan, PRESTO JST, Saitama, Japan

Kaoru Sugimura

Affiliations Institute for Integrated Cell-Material Sciences, Kyoto University, Kyoto, Japan, RIKEN Brain Science Institute, Saitama, Japan

Ruth Lehmann

Affiliation HHMI and Developmental Genetics Program, Skirball Institute of Biomolecular Medicine, Department of Cell Biology, New York University Medical Center, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS RL. Performed the experiments: PSK ADR VB IBNC SI KS. Wrote the paper: HS.