About the Authors

Yu Sato

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Nariaki Inoue

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Takashi Ishikawa

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Ryo Ishibashi

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Mayu Obata

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Hideo Aoki

Affiliation Mie Prefecture Fisheries Research Laboratory, Shima, Mie, Japan

Takashi Atsumi

Affiliation Mie Prefecture Fisheries Research Laboratory, Shima, Mie, Japan

Akira Komaru

komaru@bio.mie-u.ac.jp

Affiliation Mie University, Graduate School of Bioresources, Tsu, Mie, Japan

Competing Interests

We have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS NI TI RI TA HA AK. Performed the experiments: NI TI TA HA YS MO. Analyzed the data: YS AK. Contributed reagents/materials/analysis tools: YS MO. Wrote the paper: YS AK.