About the Authors

Henry S. Gibbons

Contributed equally to this work with: Henry S. Gibbons, Stacey M. Broomall, Lauren A. McNew, Evan W. Skowronski

henry.gibbons@us.army.mil

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Stacey M. Broomall

Contributed equally to this work with: Henry S. Gibbons, Stacey M. Broomall, Lauren A. McNew, Evan W. Skowronski

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Lauren A. McNew

Contributed equally to this work with: Henry S. Gibbons, Stacey M. Broomall, Lauren A. McNew, Evan W. Skowronski

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, Battelle Memorial Institute, Aberdeen Proving Ground, Maryland, United States of America

Hajnalka Daligault

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Carol Chapman

Affiliation Naval Medical Research Center, Biological Defense Research Directorate, Silver Spring, Maryland, United States of America

David Bruce

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Mark Karavis

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Michael Krepps

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, Excet Inc., Aberdeen Proving Ground, Maryland, United States of America

Paul A. McGregor

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, Science Applications International Corporation, Aberdeen Proving Ground, Maryland, United States of America

Charles Hong

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, Defense Threat Reduction Agency, Fort Belvoir, Virginia, United States of America

Kyong H. Park

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Arya Akmal

Affiliation Naval Medical Research Center, Biological Defense Research Directorate, Silver Spring, Maryland, United States of America

Andrew Feldman

Affiliation Johns Hopkins University Applied Physics Laboratory, Laurel, Maryland, United States of America

Jeffrey S. Lin

Affiliation Johns Hopkins University Applied Physics Laboratory, Laurel, Maryland, United States of America

Wenling E. Chang

Affiliation The MITRE Corporation, McLean, Virginia, United States of America

Brandon W. Higgs

Affiliation The MITRE Corporation, McLean, Virginia, United States of America

Plamen Demirev

Affiliation Johns Hopkins University Applied Physics Laboratory, Laurel, Maryland, United States of America

John Lindquist

Affiliation Department of Bacteriology, University of Wisconsin, Madison, Wisconsin, United States of America

Alvin Liem

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, OptiMetrics Inc, Abingdon, Maryland, United States of America

Ed Fochler

Affiliations BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America, OptiMetrics Inc, Abingdon, Maryland, United States of America

Timothy D. Read

Current address: Emory University School of Medicine, Emory University, Atlanta, Georgia, United States of America

Affiliation Naval Medical Research Center, Biological Defense Research Directorate, Silver Spring, Maryland, United States of America

Roxanne Tapia

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Shannon Johnson

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Kimberly A. Bishop-Lilly

Affiliation Naval Medical Research Center, Biological Defense Research Directorate, Silver Spring, Maryland, United States of America

Chris Detter

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Cliff Han

Affiliation Department of Energy Joint Genome Institute, Los Alamos National Laboratories, Los Alamos, New Mexico, United States of America

Shanmuga Sozhamannan

Affiliation Naval Medical Research Center, Biological Defense Research Directorate, Silver Spring, Maryland, United States of America

C. Nicole Rosenzweig

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Evan W. Skowronski

Contributed equally to this work with: Henry S. Gibbons, Stacey M. Broomall, Lauren A. McNew, Evan W. Skowronski

Current address: Operational Surveyors, Incline Village, Nevada, United States of America

Affiliation BioSciences Division, Edgewood Chemical Biological Center, Aberdeen Proving Ground, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HSG TDR EWS SS CD. Performed the experiments: SMB C. Hong CC AA HD DB RT SJ C. Han PAM LAM M. Karavis KAB-L SS. Analyzed the data: HSG LAM HD CC M. Karavis KHP AF LAM HD CC M. Krepps KHP AF JSL WEC BWH PD AL EF RT SJ KAB-L CNR. Contributed reagents/materials/analysis tools: JL. Wrote the manuscript: HSG.