About the Authors

Haitske Graveland

h.graveland@uu.nl

Affiliations Division Environmental Epidemiology, Institute for Risk Assessment Sciences, Utrecht University, Utrecht, The Netherlands, Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands, Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Jaap A. Wagenaar

Affiliations Department of Infectious Diseases and Immunology, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands, Central Veterinary Institute of Wageningen UR, Lelystad, The Netherlands

Kelly Bergs

Affiliation Division Environmental Epidemiology, Institute for Risk Assessment Sciences, Utrecht University, Utrecht, The Netherlands

Hans Heesterbeek

Affiliation Department of Farm Animal Health, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Dick Heederik

Affiliations Division Environmental Epidemiology, Institute for Risk Assessment Sciences, Utrecht University, Utrecht, The Netherlands, Julius Center for Health Sciences and Primary Care, University Medical Center, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HG JW DH HH. Performed the experiments: HG JW KB. Analyzed the data: HG DH. Contributed reagents/materials/analysis tools: HG JW DH. Wrote the paper: HG JW HH DH.