About the Authors

J. Jeffrey Morris

Affiliation Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Zackary I. Johnson

Affiliation Nicholas School of the Environment, Duke University Marine Laboratory, Beaufort, North Carolina, United States of America

Martin J. Szul

Affiliation Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Martin Keller

Affiliation Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Erik R. Zinser

ezinser@utk.edu

Affiliation Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJM ERZ. Performed the experiments: JJM MJS. Analyzed the data: JJM ZIJ MJS MK ERZ. Contributed reagents/materials/analysis tools: ZIJ MK ERZ. Wrote the paper: JJM ERZ.