About the Authors

Haruka Imada

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Masahito Hoki

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Yuji Suehiro

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Teruhiro Okuyama

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Daisuke Kurabayashi

Affiliation Department of Mechanical and Control Engineering, Tokyo Institute of Technology, Tokyo, Japan

Atsuko Shimada

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Kiyoshi Naruse

Affiliation Laboratory of BioResource, National Institute for Basic Biology, Okazaki, Aichi, Japan

Hiroyuki Takeda

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Takeo Kubo

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Hideaki Takeuchi

takeuchi@biol.s.u-tokyo.ac.jp

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HI TK HT. Performed the experiments: HI MH. Analyzed the data: HI MH HT. Contributed reagents/materials/analysis tools: HI YS TO DK AS KN HT. Wrote the paper: HI HT.