About the Authors

Yoichiro Hoshino

Affiliations Gifu Prefectural Livestock Research Institute, Takayama, Gifu, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Noboru Hayashi

Affiliations Gifu Prefectural Livestock Research Institute, Takayama, Gifu, Japan, Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Shunji Taniguchi

Affiliations Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Naohiko Kobayashi

Affiliations Gifu Prefectural Livestock Research Institute, Takayama, Gifu, Japan, Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Kenji Sakai

Affiliations Gifu Prefectural Livestock Research Institute, Takayama, Gifu, Japan, Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan

Tsuyoshi Otani

Affiliations Gifu Prefectural Livestock Research Institute, Takayama, Gifu, Japan, Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan

Akira Iritani

Affiliations Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Kazuhiro Saeki

saeki@waka.kindai.ac.jp

Affiliations Department of Genetic Engineering, Kinki University, Kinokawa Wakayama, Japan, Wakayama Industry Promotion Foundation, Wakayama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YH KS TO AI KS. Performed the experiments: YH NH ST NK KS. Wrote the paper: YH KS.